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- Fellow of the American Association for the Advancement of Science (AAAS) – 2020
- Fellow of the American Academy of Microbiology – 2009
- Arthur J. Maurer Extra Mile Award, University of Wisconsin-Madison – 2006
- Spitze Land Grant Faculty Award, UW-Madison – 2002
- Fellow, American Academy of Arts and Sciences – 2001
- American Society of Microbiology Division H Lecturer – 2001
- Vilas Associate Award, UW-Madison – 1997-1999
- Oliver H. Smith Memorial Lecturer, Marquette University – 1989
- Vilas Associate Award, University of Wisconsin-Madison – 1997
- Chairman: Department of Biochemistry, UW-Madison – 1986-1991
- George A. Miller Visiting Professor of Microbiology, Univ. of Illinois – 1986
- Evelyn Mercer Professor of Biochemistry and Molecular Biology, UW-Madison – 1985-Present
- Faculty Star Award, UW-Madison – 1984-1985
- Pasteur Award, Illinois Society for Microbiology – 1981
- Harry and Evelyn Steenbock Career Development Award, UW-Madison – 1974-1978
- NIH Career Development Award, UW-Madison – 1972-1977
- Medical Research Foundation of Boston postdoctoral fellow, Harvard Medical School – 1970
- NIH postdoctoral fellow, Harvard Medical School – 1968-1969
- NIH predoctoral fellow, Johns Hopkins University – 1963-1967
- Phi Beta Kappa, Williams College – 1963